EP2104423A1 - Verfahren zur verbesserten nutzung des produktionspotentials transgener pflanzen - Google Patents

Verfahren zur verbesserten nutzung des produktionspotentials transgener pflanzen

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Publication number
EP2104423A1
EP2104423A1 EP07856774A EP07856774A EP2104423A1 EP 2104423 A1 EP2104423 A1 EP 2104423A1 EP 07856774 A EP07856774 A EP 07856774A EP 07856774 A EP07856774 A EP 07856774A EP 2104423 A1 EP2104423 A1 EP 2104423A1
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EP
European Patent Office
Prior art keywords
spp
bacterial
fungal pathogens
plant
cryla
Prior art date
Legal status (The legal status is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the status listed.)
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Application number
EP07856774A
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German (de)
English (en)
French (fr)
Inventor
Wolfram Andersch
Reiner Fischer
Anton Kraus
Heike Hungenberg
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Bayer Intellectual Property GmbH
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Bayer CropScience AG
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Publication of EP2104423A1 publication Critical patent/EP2104423A1/de
Withdrawn legal-status Critical Current

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Classifications

    • A—HUMAN NECESSITIES
    • A01—AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
    • A01N—PRESERVATION OF BODIES OF HUMANS OR ANIMALS OR PLANTS OR PARTS THEREOF; BIOCIDES, e.g. AS DISINFECTANTS, AS PESTICIDES OR AS HERBICIDES; PEST REPELLANTS OR ATTRACTANTS; PLANT GROWTH REGULATORS
    • A01N43/00—Biocides, pest repellants or attractants, or plant growth regulators containing heterocyclic compounds
    • A01N43/34—Biocides, pest repellants or attractants, or plant growth regulators containing heterocyclic compounds having rings with one nitrogen atom as the only ring hetero atom
    • A01N43/36—Biocides, pest repellants or attractants, or plant growth regulators containing heterocyclic compounds having rings with one nitrogen atom as the only ring hetero atom five-membered rings

Definitions

  • the invention relates to a method for improving the utilization of the production potential of transgenic plants.
  • transgenic crops in agriculture has increased significantly in recent years, although regional differences are still discernible.
  • the proportion of transgenic maize in the United States has doubled from 26% to 52% since 2001, whereas transgenic maize in Germany has hardly played a practical role so far.
  • the proportion of transgenic maize is already around 12%.
  • Transgenic plants are mainly used to use the production potential of the respective plant variety as low as possible with the use of production resources as low as possible.
  • the aim of the genetic modification of the plants is primarily to produce a resistance in the plants against certain pests or harmful organisms or else herbicides and against abiotic stress (for example drought, heat or elevated salt contents).
  • a plant may be genetically modified to produce certain quality or product characteristics, such as e.g. To increase the content of selected vitamins or oils, or to improve certain fiber properties.
  • a herbicide resistance or tolerance can be achieved for example by the incorporation of genes into the crop for the expression of enzymes for detoxification of certain herbicides, which thus can grow as freely as possible, even in the presence of these herbicides for controlling weeds and grass weeds.
  • Examples include cotton varieties or corn varieties that tolerate the herbicidal active ingredient Gyphosate (Roundup®), (Roundup Ready®, Monsanto) or the herbicides glufosinate or oxynil.
  • crops that contain two or more genetic alterations have been developed, for example Monsanto has developed multi-transgenic maize varieties that target the European corn borer ⁇ Ost ⁇ nia nubilalis) and the Also known are corn or cotton crops that are resistant to both the Western corn rootworm and the cotton bollworm, as well as the herbicide Roundup®. It has now been found that the use of the production potential of transgenic crops can be further improved if the plants are treated with one or more 3-arylpyrrolidine-2,4-dione derivative (s). In this context, the term "treatment” includes all measures which lead to contact between these active substances and at least one plant part.
  • Plant parts are to be understood as meaning all aboveground and underground parts and organs of the plants, such as shoot, leaf, flower and root.
  • leaves, needles, stems, stems, flowers, fruiting bodies, fruits and seeds, as well as roots, tubers and rhizomes are listed.
  • the plant parts also include crops and vegetative and generative propagation material, such as cuttings, tubers, rhizomes, offshoots and seeds.
  • 3-Arylpyrrolidine-2,4-dione derivatives and their herbicidal or insecticidal activities are well known in the art.
  • EP-A-355 599 and EP-A-415 211 disclose bicyclic 3-aryl-pyrrolidine-2,4-dione derivatives.
  • EP-A-377 893 and EP-A-442 077 substituted monocyclic 3-aryl-pyrrolidine-2,4-dione derivatives are known.
  • EP-A-442 073 polycyclic 3-arylpyrrolidine-2,4-dione derivatives
  • EP-A-521 334 EP-A-596 298, EP-A -613,884, WO 95/01 997, WO 95/26 954, WO 95/20 572, EP-A-0 668 267, WO 96/25 395, WO 96/35 664, WO 97/01 535, WO 97/02 243, WO 97/36 868, WO 97/43 275, WO 98/05 638, WO 98/06 721, WO 98/25 928, WO 99/16748, WO 99/24 437, WO 99/43 649, WO 99/48869, WO 99/55 673, WO 01/09 092, WO 91/17972, WO 01/23 354, WO 01/74 770, WO 03/0
  • the 3 -APD usable according to the invention have the general formula (I) as follows:
  • X is halogen, alkyl, alkoxy, haloalkyl, haloalkoxy or cyano
  • W, Y and Z independently of one another represent hydrogen, halogen, alkyl, alkoxy, haloalkyl, haloalkoxy or cyano,
  • A is hydrogen, in each case optionally halogen-substituted alkyl, alkoxyalkyl, saturated, optionally substituted cycloalkyl in which optionally at least one ring atom is replaced by a heteroatom,
  • B is hydrogen or alkyl
  • a and B together with the carbon atom to which they are attached, represent a saturated or unsaturated unsubstituted or substituted cycle, optionally containing at least one heteroatom,
  • D is hydrogen or an optionally substituted radical from the group consisting of alkyl, alkenyl, alkoxyalkyl, saturated cycloalkyl, in which optionally one or more ring members are replaced by heteroatoms,
  • a and D together with the atoms to which they are bonded represent a saturated or unsaturated and optionally at least one heteroatom-containing, unsubstituted or substituted in the A, D-part cycle,
  • G is hydrogen (a) or one of the groups
  • E is a metal ion or an ammonium ion
  • L is oxygen or sulfur
  • M is oxygen or sulfur
  • RI is in each case optionally halogen-substituted alkyl, alkenyl, alkoxyalkyl, alkylthioalkyl, polyalkoxyalkyl or optionally halogen, alkyl or alkoxy-substituted cycloalkyl which may be interrupted by at least one heteroatom, in each case optionally substituted phenyl, phenylalkyl, hetaryl, phenoxyalkyl or hetaryloxyalkyl stands,
  • R.2 is in each case optionally halogen-substituted alkyl, alkenyl, alkoxyalkyl, polyalkoxyalkyl or in each case optionally substituted cycloalkyl, phenyl or benzyl,
  • R 3 represents optionally halogen-substituted alkyl or optionally substituted phenyl
  • R ⁇ and R ⁇ independently of one another are each optionally halogen-substituted alkyl, alkoxy, alkylamino, dialkylamino, alkylthio, alkenylthio, cycloalkylthio or in each case optionally substituted phenyl, benzyl, phenoxy or phenylthio and
  • R ⁇ and R ⁇ independently of one another represent hydrogen, in each case optionally halogen-substituted alkyl, cycloalkyl, alkenyl, alkoxy, alkoxyalkyl, optionally substituted phenyl, optionally substituted benzyl or together with the N-atom to which they are attached, optionally are interrupted by oxygen or sulfur optionally substituted ring.
  • At least one insecticidally active 3-APD derivative is used to treat transgenic crops.
  • insecticidally active or "insecticide” for the purposes of the invention includes insecticidal, acaricidal, molluscicidal, nematicidal, ovicidal effects, as well as a repellent, behavior modifier or sterilizing effect on pests.
  • Preferred insecticidal active compounds are compounds of the formula (I) in which
  • W is preferably hydrogen, C 1 -C 4 -alkyl, C 1 -C 4 -alkoxy, chlorine, bromine or fluorine,
  • X is preferably C 1 -C 4 -alkyl, C 1 -C 4 -alkoxy, C 1 -C 4 -haloalkyl, fluorine, chlorine or bromine, Y and Z independently of one another preferably represent hydrogen, C 1 -C 4 -alkyl, halogen, C 1 -C 4 -alkoxy or C 1 -C 4 -haloalkyl,
  • A is preferably hydrogen or in each case optionally halogen-substituted C 1 -C 6 -alkyl or C 3 -C 9 -cycloalkyl,
  • B is preferably hydrogen, methyl or ethyl
  • A, B and the carbon atom to which they are attached preferably represent saturated C3-C 6 - cycloalkyl in which optionally one ring member is replaced by oxygen or sulfur and which is optionally monosubstituted or disubstituted by Ci-Gj-alkyl, trifluoromethyl or C C 4 -alkoxy is substituted,
  • D is preferably hydrogen, in each case optionally fluorine- or chlorine-substituted C 1 -C 6 -alkyl, C 3 -C 4 -alkenyl or C 3 -C 6 -cycloalkyl,
  • a and D together preferably represent in each case optionally methyl-substituted C 3 -C 4 -alkanediyl, in which optionally a methylene group is replaced by sulfur,
  • G is preferably hydrogen (a) or one of the groups
  • E ⁇ or ⁇ - N ' ⁇ ⁇ 7 7 (g) in particular stands for (a), (b), (c) or (g)
  • E is a metal ion or an ammonium ion
  • M is oxygen or sulfur
  • R 1 preferably represents in each case optionally halogen-substituted Ci-Cio-alkyl, C 2 -C 0 - alkenyl, Ci-C4-alkoxy-Ci-C4-alkyl, Ci-C4-alkylthio-Ci-C 4 alkyl or C 3 -C 6 -cycloalkyl optionally substituted by fluorine, chlorine, C 1 -C 4 -alkyl or C 1 -C 2 -alkoxy, optionally substituted by fluorine, chlorine, bromine, cyano, nitro, C 1 -C 4 -alkyl, C r is C 4 -alkoxy, Trifluoromethyl or trifluoromethoxy-substituted phenyl, in each case optionally substituted by chlorine or methyl-substituted pyridyl or thienyl,
  • R 2 preferably represents in each case optionally substituted by fluorine or chlorine, Ci-Cio-alkyl, C 2 -C O alkenyl, Ci-C 4 -alkoxy-C 2 -C 4 alkyl, optionally substituted by methyl or methoxy-substituted C 5 - C 6 -cycloalkyl or is in each case optionally substituted by fluorine, chlorine, bromine, cyano, nitro, C 1 -C 4 -alkyl, C 1 -C 4 -alkoxy, trifluoromethyl or trifluoromethoxy-substituted phenyl or benzyl,
  • R 3 preferably represents optionally fluorine-substituted C 1 -C 4 -alkyl or each optionally substituted by fluorine, chlorine, bromine, C 1 -C 4 -alkyl, C 1 -C 4 -alkoxy, trifluoromethyl, trifluoromethoxy, cyano or nitro-substituted phenyl .
  • R 4 preferably represents in each case optionally substituted by fluorine or chlorine, C r C 4 alkyl, Ci-C 4 alkoxy, Ci-C 4 alkylamino, Ci-C4-alkylthio, or represents in each case optionally substituted by fluorine, chlorine, bromine, nitro , cyano, Ci-C 4 alkoxy, trifluoromethoxy, Ci-C 4 alkylthio, Q-C4 haloalkylthio, Ci-C 4 -alkyl or trifluoromethyl-substituted phenyl, phenoxy or phenylthio,
  • R 5 is preferably C 1 -C 4 -alkoxy or C 1 -C 4 -thioalkyl
  • R 6 preferably represents C r C 6 alkyl, C 3 -C 6 cycloalkyl, C, -C 6 alkoxy, C 3 -C 6 alkenyl, QC 4 - alkoxy-C r C 4 -alkyl,
  • R 7 preferably represents QC 6 alkyl, C 3 -C 6 alkenyl or C r C 4 alkoxy-C r C 4 alkyl,
  • W particularly preferably represents hydrogen, methyl, ethyl, chlorine, bromine or methoxy
  • X particularly preferably represents chlorine, bromine, methyl, ethyl, propyl, i-propyl, methoxy, ethoxy or trifluoromethyl,
  • Y and Z more preferably independently of one another represent hydrogen, fluorine, chlorine, bromine, methyl, ethyl, propyl, isopropyl, trifluoromethyl or methoxy
  • B particularly preferably represents hydrogen, methyl or ethyl
  • A, B and the carbon atom to which they are attached are particularly preferably saturated C ⁇ -cycloalkyl, in which optionally a ring member is replaced by oxygen and which is optionally monosubstituted by methyl, ethyl, methoxy, ethoxy, propoxy or butoxy,
  • D is particularly preferably hydrogen, methyl, ethyl, propyl, isopropyl, butyl, isobutyl, allyl, cyclopropyl, cyclopentyl or cyclohexyl,
  • a and D together are particularly preferred for optionally methyl-substituted C 3 -C 4 -alkanediyl
  • G is particularly preferably hydrogen (a) or one of the groups
  • M is oxygen or sulfur
  • R 1 particularly preferably represents C 1 -C 6 -alkyl, C 2 -C 4 -alkenyl, methoxymethyl, ethoxymethyl, ethylthiomethyl, cyclopropyl, cyclopentyl or cyclohexyl,
  • R 2 particularly preferably represents C 1 -C 6 -alkyl, C 2 -C 4 -alkenyl, methoxyethyl, ethoxyethyl or represents phenyl or benzyl, R 6 and R 7 independently of one another particularly preferably represent methyl, ethyl or together represent a C 5 -alkylene radical in which the C 3 -methylene group is replaced by oxygen.
  • W very particularly preferably represents hydrogen or methyl
  • X very particularly preferably represents chlorine, bromine or methyl
  • Y and Z very particularly preferably independently of one another represent hydrogen, chlorine, bromine or methyl
  • A, B and the carbon atom to which they are attached are very particularly preferably saturated C 6 -cycloalkyl, in which optionally a ring member is replaced by oxygen and which is optionally monosubstituted by methyl, trifluoromethyl, methoxy, ethoxy, propoxy or butoxy .
  • G is very particularly preferably hydrogen (a) or one of the groups
  • M is oxygen or sulfur
  • R 1 very particularly preferably for Cj-Cg-alkyl, C 2 -C 4 - alkenyl, methoxymethyl, ethoxymethyl, ethylthiomethyl, cyclopropyl, cyclopentyl, cyclohexyl or
  • R 2 very particularly preferably represents C 1 -C 8 -alkyl, C 2 -C 4 -alkenyl, methoxyethyl, ethoxyethyl, phenyl or benzyl, R 6 and R 7 independently of one another very particularly preferably represent methyl, ethyl or together represent a Cs-alkylene radical in which the C 3 -methylene group is replaced by oxygen.
  • the compounds of the formula (I) can also be present as optical isomers or mixtures of isomers in different compositions.
  • the compounds of formula (I) are - as mentioned above - known in the art; as well as their preparation (see in particular WO 97/01 535, WO 97/36 868, WO 98/05 638, WO 04/007 448).
  • transgenic plants in particular crops, are treated with 3-APD derivatives in order to increase agricultural productivity.
  • transgenic plants are plants which code at least one gene or gene fragment which they have not taken over by fertilization.
  • This gene or gene fragment may be derived or derived ("foreign gene") from another plant of the same species, from plants of different species, but also from organisms from the animal kingdom or microorganisms (including viruses) and / or may already be natural
  • the use of synthetic genes is also possible according to the invention and is also subsumed here under the term "foreign gene”. It is also possible that a transgenic plant encodes two or more foreign genes of different origin.
  • the "foreign gene” in the sense of the invention is furthermore characterized in that it comprises a nucleic acid sequence which exerts a specific biological or chemical function or activity in the transgenic plant .
  • These genes generally encode biocatalysts, such as, for example, enzymes or ribozymes. or include regulatory sequences such as promoters or terminators for influencing the expression of endogenous proteins, but may also encode regulatory proteins such as repressors or inducers, and the foreign gene may also serve the targeted localization of a gene product of the transgenic plant, for example Also inhibitors such as antisense RNA can be encoded by the foreign gene.
  • GURT technology Genetic Use Restriction Technologies
  • two or three foreign genes are cloned into the crop which, in a complex interplay following the delivery of an external stimulus, trigger a cascade that results in the death of the otherwise developing embryo, for example, the external stimulus (eg, an agent or another chemical or abiotic stimulus) may interact with a repressor thus no longer suppress expression of a recombinase so that the recombinase can cleave an inhibitor and thus allow the expression of a toxin causing the embryo to die off
  • the external stimulus eg, an agent or another chemical or abiotic stimulus
  • transgenic plants which have altered properties due to the integration of regulatory foreign genes and the optionally mediated overexpression, suppression or inhibition of endogenous genes or gene sequences or the existence or expression of foreign genes or their fragments.
  • the method according to the invention makes it possible to improve the utilization of the production potential of transgenic plants.
  • this may possibly be attributable to the fact that the amount of Aurwand of the active substance which can be used according to the invention can be reduced; for example, by reducing the dose used or by reducing the number of applications.
  • the yield of crops can be increased quantitatively and / or qualitatively. This is especially true in a transgenic resistance to biotic or abiotic stress.
  • insecticidal 3-APD is used, the dosage of the insecticide may u. be limited to a sub-lethal dose without significantly attenuating the desired effect of the active ingredient on the pests.
  • synergistic effects can vary depending on the plant species or plant cultivars, their location and growth conditions (soils, climate, vegetation period, diet) and can be diverse.
  • reduced application rates and / or extensions of the spectrum of action and / or an increase in the effect of the substances and agents usable in the invention better plant growth, increased tolerance to high or low temperatures, increased tolerance to drought or to water or soil salt content, increased flowering power facilitated harvest, acceleration of ripeness, higher crop yields, higher quality and / or higher nutritional value of the harvested products, higher shelf life and / or machinability of the harvested products, which exceed the actual expected effects.
  • transgenic plants A list of examples of transgenic plants known to the person skilled in the art, with reference in each case to the structure in question in the plant or in the plant expressed by the genetic modification, is compiled in Table 1. This is the affected one Structure or the expressed principle in each case assigned a specific feature expression in the sense of a tolerance to a certain stress factor.
  • Table 3 A similar list (Table 3) also provides, in somewhat different order, examples of modes of action, induced tolerances and potential crops. Further examples of transgenic plants which are suitable for the treatment according to the invention are compiled in Tables 4 and 5 and 6.
  • the 3-APD are used for the treatment of transgenic plants containing at least one gene or gene fragment for coding a Bt toxin.
  • a Bt toxin is a protein originally derived or derived from the soil bacterium, Bacillus thuringiensis, which belongs to either the group of crystal toxins (Cry) or cytolytic toxins (Cyt). They are originally formed in the bacterium as protoxins and are only in an alkaline environment - for example, in the digestive tract of certain feeding insects - metabolized to its active form. There, the then active toxin binds to certain hydrocarbon structures on cell surfaces and causes pore formation, which can destroy the osmotic potential of the cell and thus cause cell lysis. The death of insects is the result.
  • Bt toxins are effective against certain pest species of the order Lipidoptera (butterflies), Homoptera, Diptera (fly) and Coleoptera (beetles) in all their stages of development; from the Eilarve to her juvenile to her adult forms.
  • Bt plants It has long been known to clone gene sequences encoding Bt toxins, parts thereof, or Bt toxin-derived proteins or peptides into agricultural crops using genetic engineering techniques, and thus to generate transgenic plants with endogenous resistance to Bt toxin-susceptible pests.
  • the transgenic plants encoding at least one Bt toxin or proteins derived therefrom are defined as "Bt plants" for the purposes of the invention.
  • the "first generation" of such Bt plants usually contained only the genes that allowed the formation of a specific toxin and thus made it resistant only to a group of pests, examples of a commercially available maize strain containing the gene for the formation of CrylAb Toxins is "YieldGard®” from Monsanto, which is resistant to the European corn borer.
  • the resistance against other pest pathogens of the family of Lepidoptera is generated in the Bt cotton variety (Bollgard®) by cloning the genes to form the CrylAc toxin.
  • Still other transgenic crop plants express genes for the production of Bt toxins with action against pathogens from the order of coleopterans.
  • Examples include the Bt potato variety “NewLeaf®” (Monsanto), which can form the Cry3A toxin and is thus resistant to the Colorado beetle ("Colorado Potatoe Beeile"), or the transgenic maize variety “YieldGard®” (Monsanto), which form the Cry 3BbI toxin and thus protected against various species of corn rootworm.
  • transgenic plants with Bt toxins from the group of the Cry family (see, for example, Crickmore et al., 1998, Microbiol. Mol. Biol. Rev. 62: 807-812), which are particularly effective against Lepidoptera, Coleoptera or Diptera.
  • Bt toxins from the group of the Cry family (see, for example, Crickmore et al., 1998, Microbiol. Mol. Biol. Rev. 62: 807-812), which are particularly effective against Lepidoptera, Coleoptera or Diptera.
  • genes encoding the proteins are:
  • crylAal crylAa2, crylAa3, crylAa4, crylAa5, crylAa ⁇ , crylAa7, crylAa ⁇ , crylAa9, crylAalO, crylAal 1crylAbl, crylAb2, crylAb3, crylAb4, crylAb5, crylAb ⁇ , crylAb7, crylAb ⁇ , crylAb9, crylAblO, crylAbl l, crylAbl2, crylAbl3, crylAbH , crylAcl, crylAc2, crylAc3, crylAc4, crylAc5, crylAc ⁇ , crylAc7, crylAc ⁇ , crylAc9, crylAclO, crylAcl1, crylAcl2, crylAcl3, crylAdl, crylAd2, crylAel, crylAfl, crylAgl, crylBal, crylBa2, crylBbl, crylBcl
  • plants which, in addition to the genes for one or more Bt toxins, may also contain genes for expression of, for example, a protease or peptidase inhibitor (as in WO-A 95/35031), herbicidal resistance (eg Glufosinate or glyphosate by expression of the pat or bar gene) or to the formation of nematode, fungal or viral resistances (eg by expression of a glucanase, chitinase) contain or express.
  • a protease or peptidase inhibitor as in WO-A 95/35031
  • herbicidal resistance eg Glufosinate or glyphosate by expression of the pat or bar gene
  • fungal or viral resistances eg by expression of a glucanase, chitinase
  • ingredients may also be genetically modified in their metabolic properties so as to exhibit a qualitative and / or quantitative modification of ingredients (e.g., by modifications of energy, carbohydrate, fatty acid, or nitrogen metabolism or affecting metabolite fluxes) (see above).
  • Table 2 A list of examples of principles of action which can be introduced into a crop by genetic modification and which are suitable for the treatment according to the invention, alone or in combination, is shown in Table 2. This table contains under the indication "AP" (active principle) the respective active principle and the associated pest to be controlled.
  • AP active principle
  • the process according to the invention is used for the treatment of transgenic vegetable, maize, soybean, cotton, tobacco, rice, potato and sugar beet varieties. These are preferably Bt plants.
  • the vegetables or varieties are, for example, the following crops:
  • o Potatoes preferably starch potatoes, sweet potatoes and ware potatoes;
  • o root vegetables preferably carrots, turnips (turnips, turnips, turnips, turnips, Teltower turnips), salsify, Jerusalem artichoke, root parsley, parsnip, radish and horseradish;
  • o Tubers preferably kohlrabi, beetroot, celeriac, radishes and radishes;
  • o onion vegetables preferably leek, leek and onions (poplar and seedling);
  • cabbage preferably cabbage (white cabbage, red cabbage, kale, savoy cabbage), cauliflower, sprouts, broccoli, kale, marrow cabbage, sea cabbage and Brussels sprouts; o Fruit vegetables: preferably tomatoes (field, shrub, meat, greenhouse, cocktail, industrial and fresh market tomatoes), melons, eggplant, aubergines, peppers (vegetable and spice paprika, Spanish pepper), hot peppers, pumpkin, Zucchini and cucumbers (field, greenhouse, snake and cucumbers);
  • Vegetable legumes preferably wild beans (as sword beans, pea beans, flageolet beans, wax beans, dry beans with green and yellow-hulled varieties), runner beans (as sword beans, pea beans, flageolet beans, green beans with green, blue and yellow-hulled varieties), broad beans (broad beans, Broad beans, varieties with white and black spotted flowers), peas (chick peas, chickpeas, peas, raspberries, sugar peas, pea peas, varieties with light and dark green fresh grain) and lentils;
  • Leaf and stalk vegetables preferably Chinese cabbage, lettuce, pickle salad, corn salad, iceberg lettuce, romaine lettuce, le weree salad, endives, radicchio, lollo rosso salad, rocket salad, chicory, spinach, Swiss chard and parsley;
  • Bt vegetables including exemplary methods for their production, are described in detail, for example, in Barton et al., 1987, Plant Physiol. 85: 1103-1109; Vaeck et al., 1987, Nature 328: 33-37; Fischhoff et al., 1987, Bio / Technology 5: 807-813.
  • Bt vegetable plants are already known as commercially available variants such as the potato variety NewLeaf ® (Monsanto) beyond.
  • US 6,072,105 describes the production of Bt vegetables.
  • Bt cotton is already known in principle, for example from US Pat. No. 5,322,938 or from Prietro-Samson ⁇ r et al., J. Ind. Microbiol. & Biotechn. 1997, 19, 202, and H. Agaisse and D. Lereclus, J. Bacteriol. 1996, 177, 6027.
  • Bt cotton is already commercially available in various variations, for example under the name NuCOTN® (Deltapine (USA)).
  • NuCOTN® Deltapine (USA)
  • the Bt cotton NuCOTN33® and NuCOTN33B® is particularly preferred.
  • Bt maize has also been known for a long time, for example from Ishida, Y., Saito, H., Ohta, S., Hiei, Y., Komari, T., and Kumashiro, T. (1996).
  • High efficiency transformation of maize Zea mayz L.
  • EP-B-0485506 also describes the production of Bt maize plants.
  • Bt maize is also commercially available in various variations, for example under the following names (company / companies in brackets): KnockOut® (Novartis Seeds), SkypeGard® (Mycogen Seeds), Yieldgard® (Novartis Seeds, Monsanto, Cargill, Golden Harvest , Pioneer, DeKalb and others), Bt-Xtra® (DeKalb) and StarLink® (Aventis CropScience, Garst et al.).
  • the following maize varieties are particularly preferred: KnockOut®, NaturGard®, Yieldgard®, Bt-Xtra® and StarLink®.
  • soybean Roundup®Ready varieties or varieties are available with a resistance to the herbicide Liberty Link® and treatable according to the invention.
  • a variety of lines of the so-called “Golden Rice” are available, which are equally characterized by having an increased content of provitamin A by a transgenic modification.Also here are examples of plants that are after the invention Process with the described advantages can be treated.
  • the method according to the invention is suitable for controlling a multiplicity of harmful organisms which occur in particular in vegetables, maize or cotton, in particular insects and arachnids, very particularly preferably insects.
  • the mentioned pests include:
  • Thysanura for example, Lepisma spp.
  • Orthoptera for example Blattella spp., Periplaneta spp., Leucophaea spp., Acheta spp., Gryllotalpa spp., Locusta migratoria migratorioides, Melanoplus spp., Schistocerca spp.
  • Thysanoptera for example Frankliniella spp., Kakothrips spp., Hercinothrips spp., Scirtothrips spp., Taeniothrips spp., Thrips spp.
  • Lepidoptera from the order of Lepidoptera: for example, Pectinophora spp., Bupalus spp., Cheimatobia spp., Cnephasia spp., Hydraecia spp., Lithocolletis spp., Hyponomeuta spp., Plutella spp., Plutella xylostella, Malacosoma spp., Euproctis spp.
  • Anobium spp. Rhizopertha spp., Bruchidius spp., Acanthoscelides spp., Hylotrupes spp., Aclypea spp., Agelastica spp., Leptinotarsa spp., Psylliodes spp., Chaetocnema spp., Cassida Spp., Bothynoderes spp., Clivina spp., Ceutorhynchus spp., Phyllotreta spp., Apion spp., Sitona spp., Bruchus spp., Phaedon spp., Diabrotica spp., Psylloides spp., Epilachna spp., Atomaria spp.
  • Oryzaephilus spp. Anthonomus spp., Sitophilus spp., Otiorhynchus spp., Cosmopolites spp., Ceuthorrynchus spp., Hypera spp., Dermestes spp., Trogoderma spp., Anthrenus spp., Attagenus spp., Lyctus spp., Meligethes Spp., Ptinus spp., Niptus spp., Gibbium spp., Tribolium spp., Tenebrio spp., Agriotes spp., Conoderus spp., Melolontha spp., Amphimallon spp., Costelytra spp. o from the order Hymenoptera: for example Diprion spp., Hoplocampa spp., Lasius spp., Monomorium spp., Ve
  • Drosophila spp. for example Drosophila spp., Chrysomyxa spp., Hypoderma spp., Tannia spp., Bibio spp., Oscinella spp., Phorbia spp., Pegomyia spp., Ceratitis spp., Dacus spp., Tipula spp ..
  • Acarina for example Acarus spp., Bryobia spp., Panonychus spp., Tetranychus spp., Eotetranychus spp., Oligonychus spp., Eutetranychus spp., Eriophyes spp., Phyllocoptruta spp., Tarsonemus spp., Rhipicephalus spp., Rhipicephalus spp., Ixodes spp., Amblyomma spp.
  • helminth class for example Meloidogyne spp., Heterodera spp., Globodera spp., Radopholus spp., Pratylenchus spp., Tylenchulus spp., Tylenchorhynchus spp., Rotylenchus spp., Heliocotylenchus spp., Belonoaimus spp., Longidorus Spp., Trichodorus spp., Xiphinema spp., Ditylenchus spp., Aphelenchoides spp., Anguina spp.
  • Meloidogyne spp. for example Meloidogyne spp., Heterodera spp., Globodera spp., Radopholus spp., Pratylenchus spp., Tylenchulus spp., Tylenchorhynchus spp
  • Gastropoda for example, Deroceras spp., Arion spp., Lymnaea spp., Galba spp., Succinea spp.
  • o Hemiptera order for example, species of suborder Sternorrhyncha
  • the inventive method for controlling Agriotes spp. Melolontha spp., Aphis spp., Cnephasia spp., Ostrinia spp., Agrotis spp., Hydraecia spp., Tipula spp., Myzus spp., Bemisia spp., Trialeurodes Spp., Oscinella spp., Tetranychus spp., Lygus spp., Leptinotarsa spp., Psylliodes spp., Phytometra spp., Deroceras spp., Psylla spp., Blaniulus spp., Onychiurus spp., Piesma spp., Atomaria spp.
  • the active compounds which can be used according to the invention are particularly suitable for controlling insects from the suborder of the plant lice (Sternorrhyncha), in particular for controlling the bladder lice, the root lice, the leaf flea, the Napfschildläuse, the Lupus louse, the Röhrenschildläuse or the lice and lice. This use is described in detail in WO 2006/077071 and is hereby referred to for disclosure purposes.
  • the method according to the invention for the treatment of Bt vegetables, Bt maize, Bt cotton, Bt soya, Bt tobacco as well as Bt rice, Bt sugar beets or Bt potatoes is suitable for controlling aphids (Aphidina), whites Flies (Trialeurodes), Thrips (Thysanoptera), Spider mites (Arachnida), Shags or Lice (Coccoidae or Pseudococcoidae).
  • the active compounds which can be used according to the invention can be employed in customary formulations, such as solutions, emulsions, wettable powders, water- and oil-based suspensions, powders, dusts, pastes, soluble powders, soluble granules, scattering granules, suspension-emulsion concentrates, active substance-impregnated natural products, active ingredient Impregnated synthetic materials, fertilizers and Feinstverkapselitch in polymeric materials.
  • formulations are prepared in a known manner, e.g. by mixing the active compounds with extenders, ie liquid solvents and / or solid carriers, if appropriate using surface-active agents, ie emulsifiers and / or dispersants and / or foam-forming agents.
  • extenders ie liquid solvents and / or solid carriers
  • surface-active agents ie emulsifiers and / or dispersants and / or foam-forming agents.
  • Injectable powders are preparations which are uniformly dispersible in water and which, in addition to the active substance except a diluent or inert substance, also contain wetting agents, e.g. polyoxethylated alkylphenols, polyoxethylated fatty alcohols, alkyl or alkylphenol sulfonates and dispersants, e.g. sodium lignosulfonate, 2,2'-dinaphthylmethane-6,6'-disulfonic acid sodium.
  • wetting agents e.g. polyoxethylated alkylphenols, polyoxethylated fatty alcohols, alkyl or alkylphenol sulfonates and dispersants, e.g. sodium lignosulfonate, 2,2'-dinaphthylmethane-6,6'-disulfonic acid sodium.
  • Dusts are obtained by grinding the active substance with finely divided solids, for example talc, natural clays such as kaolin, bentonite, pyrophillite or diatomaceous earth.
  • Granules can either be prepared by spraying the active ingredient onto adsorptive, granulated inert material or by applying active substance concentrates by means of Adhesives, for example polyvinyl alcohol, polyacrylic acid sodium or mineral oils, on the surface of carriers such as sand, kaolinites or granulated inert material. It is also possible to granulate suitable active ingredients in the manner customary for the production of fertilizer granules, if desired in admixture with fertilizers.
  • Adhesives for example polyvinyl alcohol, polyacrylic acid sodium or mineral oils
  • Excipients which can be used are those which are suitable for imparting special properties to the composition itself and / or preparations derived therefrom (for example spray liquor, seed dressing), such as certain technical properties and / or specific biological properties.
  • Typical auxiliaries are: extenders, solvents and carriers.
  • polar and non-polar organic chemical liquids e.g. from the classes of aromatic and non-aromatic hydrocarbons (such as paraffins, alkylbenzenes, alkylnaphthalenes, chlorobenzenes), alcohols and polyols (which may also be substituted, etherified and / or esterified), ketones (such as acetone, cyclohexanone), Esters (including fats and oils) and (poly) ethers, simple and substituted amines, amides, lactams (such as N-alkylpyrrolidones) and lactones, sulfones and sulfoxides (such as dimethyl sulfoxide).
  • aromatic and non-aromatic hydrocarbons such as paraffins, alkylbenzenes, alkylnaphthalenes, chlorobenzenes
  • alcohols and polyols which may also be substituted, etherified and / or esterified
  • ketones such
  • Suitable liquid solvents are essentially: aromatics, such as xylene, toluene, or alkylnaphthalenes, chlorinated aromatics and chlorinated aliphatic hydrocarbons, such as chlorobenzenes, chloroethylenes or methylene chloride, aliphatic hydrocarbons, such as cyclohexane or paraffins, e.g.
  • Petroleum fractions mineral and vegetable oils, alcohols such as butanol or glycol and their ethers and esters, ketones such as acetone, methyl ethyl ketone, methyl isobutyl ketone or cyclohexanone, strongly polar solvents such as dimethyl sulfoxide, and water.
  • Suitable solid carriers are, for example:
  • ammonium salts and ground natural minerals such as kaolins, clays, talc, chalk, quartz, attapulgite, montmorillonite or diatomaceous earth and synthetic minerals, such as finely divided silica, alumina and silicates, as solid carriers for granules in question: eg broken and fractionated natural rocks such Calcite, marble, pumice, sepiolite, dolomite and synthetic granules of inorganic and organic flours and granules of organic material such as paper, sawdust, coconut shells, corn cobs and tobacco stalks; as emulsifying and / or foam-producing agents come into question: eg nonionic and anionic emulsifiers, such as polyoxyethylene fatty acid esters, polyoxyethylene fatty alcohol ethers, for example alkylaryl polyglycol ethers, alkylsulfonates, alkyl sulfates, arylsulfonates and protein hydrolys
  • oligo- or polymers for example starting from vinylic monomers, from acrylic acid, from EO and / or PO alone or in combination with, for example, (poly) alcohols or (poly) amines.
  • lignin and its sulfonic acid derivatives simple and modified celluloses, aromatic and / or aliphatic sulfonic acids and their adducts with formaldehyde.
  • Adhesives such as carboxymethylcellulose, natural and synthetic powdery, granular or latex-type polymers such as gum arabic, polyvinyl alcohol, polyvinyl acetate, and natural phospholipids such as cephalins and lecithins and synthetic phospholipids may be used in the formulations.
  • Dyes such as inorganic pigments, e.g. Iron oxide, titanium oxide, ferrocyan blue and organic dyes such as alizarin, azo and metal phthalocyanine dyes and trace nutrients such as salts of iron, manganese, boron, copper, cobalt, molybdenum and zinc.
  • inorganic pigments e.g. Iron oxide, titanium oxide, ferrocyan blue and organic dyes such as alizarin, azo and metal phthalocyanine dyes and trace nutrients such as salts of iron, manganese, boron, copper, cobalt, molybdenum and zinc.
  • additives may be fragrances, mineral or vegetable optionally modified oils, waxes and nutrients (also trace nutrients), such as salts of iron, manganese, boron, copper, cobalt, molybdenum and zinc.
  • Stabilizers such as cold stabilizers, preservatives, antioxidants, light stabilizers or other chemical and / or physical stability-improving agents may also be present.
  • the plants or plant parts are treated according to the invention with an oil-based suspension concentrate.
  • An advantageous suspension concentrate is known from WO 2005/084435 (EP 1 725 104 A2). It consists of at least one agrochemically active substance solid at room temperature, at least one "closed" penetrating agent, at least one vegetable oil or mineral oil, at least one nonionic surfactant and / or at least one anionic surfactant and optionally one or more additives from the groups of emulsifying agents Foam-inhibiting agents, preservatives, antioxidants, dyes and / or inert fillers Preferred embodiments of the suspension concentrate are described in WO 2005/084435. Corresponding suspension concentrates on a vegetable oil basis are expressly described in EP 1 725 105 A2 for the present invention 3-APD, both of which are fully incorporated by reference for purposes of the disclosure.
  • the plants or the plant part are treated with agents containing ammonium or phosphonium salts and optionally penetration promoters according to the invention.
  • An advantageous agent is known from DE-05059469. It consists of at least one active substance from the class of 3-APD and at least one ammonium or phosphonium salt, if appropriate, penetration promoters. Preferred embodiments are described in DE-05059469. This document is incorporated by reference for purposes of disclosure.
  • the formulations generally contain between 0.01 and 98% by weight of active ingredient, preferably between 0.5 and 90%.
  • the active ingredient concentration is for example about 10 to 90 wt .-%, the remainder to 100 wt .-% consists of conventional formulation components.
  • the active ingredient concentration may be about 5 to 80% by weight.
  • Dust-form formulations usually contain 5 to 20% by weight of active ingredient, sprayable solutions about 2 to 20% by weight.
  • Part of active ingredient content depends on whether the active compound is liquid or solid and which granulation aids, fillers, etc. are used.
  • the active compounds according to the invention can be present in their commercially available formulations and in the use forms prepared from these formulations in mixtures with other active ingredients, such as insecticides, attractants, sterilants, acaricides, nematicides, fungicides, growth-regulating substances or herbicides.
  • Particularly favorable mixing partners are e.g. the following:
  • Inhibitors of the respiratory chain complex HI Azoxystrobin, Cyazofamide, Dimoxystrobin, Enestrobin, Famoxadone, Fenamidon, Fluoxastrobin, Kresoximmethyl, Metominostrobin, Orysastrobin, Pyraclostrobin, Picoxystrobin, Trifloxystrobin
  • Methoxy-3-pyridinyl) -cyclopropane carboxamide 1 - [(4-methoxyphenoxy) methyl] -2,2-dimethylpropyl-1H-imidazole-1-carboxylic acid, O- [1 - [(4-methoxyphenoxy) methyl] - 2,2-dimethylpropyl] -1H-imidazole-1-carbothioic acid, 2- (2 - ⁇ [6- (3-chloro-2-methylphenoxy) -5-fluoropyrimidin-4-yl] oxy ⁇ phenyl) -2- (methoxyin ⁇ ino) -N-methylacetaniid bactericides:
  • Carbamates for example alanycarb, aldicarb, aldoxycarb, allyxycarb, aminocarb, bendodi carb, benfuracarb, bufencarb, butacarb, butocarboxime, butoxycarboxim, carbarnyl, carbofuran, carbosulfan, cloethocarb, dimetilane, ethiofencarb, fenobutarb, fenothiocarb, formetanates, Furathiocarb, isoprocarb, metam-sodium, methiocarb, methomyl, metolcarb, oxamyl, pirimicarb, promecarb, propoxur, thiodicarb, thiofanox, trimethacarb, XMC, xylylcarb, triazamates
  • Organophosphates for example acephates, azamethiphos, azinphos (-methyl, -ethyl), bromophosethyl, bromfenvinfos (-methyl), butathiofos, cadusafos, carbophenothione, chloroethoxyfos, chlorfenvinphos, chlormephos, chlo ⁇ yrifos (-methyl / -ethyl), coumaphos, Cyanofenphos, cyanophos, chlorfenvinphos, demeton-S-methyl, demeton-S-methylsulphone, dialifos, diazinon, dichlofenthione,
  • Sodium Channel Modulators / Voltage-Dependent Sodium Channel Blockers Pyrethroids, for example, acrinathrin, allethrin (d-cis-trans, d-trans), beta-cyfluthrin, biferthrin, bioallethrin, bioallethrin-S-cyclopentyl isomer, bioethanomethrin, biopermethrin, bioresmethrin, chlovaporthrin, cis-cyclopenthrin , Cis-resmethrin, cis-permethrin, clocthrin, cycloprothrin, cyfluthrin, cyhalothrin, cyclopenthrin (alpha, beta, theta, zeta), cyphenothrin, deltamethrin, empenthrin (IR isomer), esfenvalerate, etofenprox, fen
  • Oxadiazines for example Indoxacarb
  • Chloronicotinyls for example acetamiprid, clothianidin, dinotefuran, imidacloprid, nitenpyram,
  • Acetylcholine receptor modulators are Acetylcholine receptor modulators
  • GABA-driven chloride channel antagonists Organochlorines, for example camphechlor, chlordane, endosulfan, gamma-HCH, HCH, heptachlor, lindane, methoxychlor
  • Fiproles for example, acetoprole, ethiprole, fipronil, pyrafluprole, pyriprole, vaniliprole
  • Mectins for example Abamectin, Emamectin, Emamectin benzoate, Ive ⁇ nectin,
  • Juvenile hormone mimetics for example, diofenolan, epofenonans, fenoxycarb, hydroprene, kinoprene,
  • Diacylhydrazines for example chromafenozide, Halofenozide, Methoxyfenozide, Tebufenozide
  • Benzoylureas for example bistrifluron, chlorofluazuron, diflubenzuron, fluazuron, flucuroxuron, flufenoxuron, hexaflumuron, lufenuron, novaluron, noviflumuron, penfluron, teflubenzuron, triflumuron
  • Organotin compounds for example azocyclotin, cyhexatin, fenbutatin oxides
  • Dinitrophenols for example binapacyrl, dinobutone, dinocap, DNOC, meptyldinocap
  • METI's for example fenazaquin, fenpyroximate, pyrimidifen, pyridaben, tebufenpyrad, tolfenpyrad
  • spirodiclofen for example spirodiclofen, spiromesifen,
  • rynaxypyr (3-bromo-N- ⁇ 4-chloro-2-methyl-6-
  • Fumigants for example aluminum phosphides, methyl bromides, sulfuryl fluorides
  • Food inhibitors for example Cryolite, Flonicamid, Pymetrozine
  • Mite growth inhibitors for example clofentezine, etoxazole, hexythiazox
  • a mixture with other known active ingredients, such as herbicides, fertilizers, growth regulators, safeners, semiochemicals, or with agents for improving the plant properties is possible.
  • the active ingredient content of the use forms prepared from the commercially available formulations may range from 0.00000001 to 95% by weight, preferably between 0.00001 and 1% by weight of active ingredient.
  • Acetolactate synthase (ALS) sulfonylurea compounds imidazolinones
  • Triazole pyrimidines pyrimidyloxybenzoates
  • Acetyl-CoA-carboxylase (ACCase) aryloxyphenoxyalkanecarboxylic acid
  • HPPD Hydroxyphenylpyruvate dioxygenase
  • Trione like mesotrione or sulcotrione
  • ADSL Adenylosuccinate lyase
  • EPSPS Synthase
  • Protoporphyrinogen oxidase diphenyl ether, cyclic imide
  • Phenopylate oxadiazole, etc.
  • Cytochrome P450 e.g. P450 SUl Xenobiotics and herbicides like
  • Rhopalosiphum maydis, Diplodia maydis, Ostrinia nubilalis, Lepidoptera sp.
  • CMi ⁇ small basic peptide building block plant pathogens, for example Fusarium, Alternaria, from corn kernel
  • Com-SAFP (Zeamatin) plant pathogens, e.g. Fusarium,
  • Envelope proteins viruses such as the corn dwarfing virus
  • Bacillus cereus toxin, photorabdus and nematodes e.g. Ostrinia nubilalis,
  • Toxins of Xenorhabdus Heliothis zea, armyworms e.g.
  • Sesamia sp. Aprotis ipsilon, Asian corn borer, weevil
  • Aminopeptidase inhibitors e.g. Leucine Lepidoptera, Coleoptera, Diptera,
  • LAPI Aminopeptidase Inhibitors
  • Heliothis zea armyworms e.g. Spodoptera frugiperda, corn rootworm, Sesamia sp.,
  • Lectin Lepidoptera Coleoptera, Diptera, Nematodes, e.g. Ostrinia nubilalis, Heliothis zea,
  • Protease inhibitors eg cystatin, patatin, weevil, corn rootworm Affected structure or expressed principle Characteristic of the plant / tolerance
  • Virgiferin CPTI Ribosome Inactivating Protein Lepidoptera, Coleoptera, Diptera, Nematodes, e.g. Ostrinia nubilalis, Heliotbis zea, armyworms e.g. Spodoptera frugiperda,
  • 5C9 maize polypeptide Lepidoptera, Coleoptera, Diptera, Nematodes, e.g. Ostrinia nubilalis, Heliotbis zea, armyworms e.g. Spodoptera frugiperda,
  • Acetolactate synthase (ALS) sulfonylurea compounds imidazolinones
  • Triazole pyrimidines pyrimidyloxybenzoates
  • Acetyl-CoA-carboxylase (ACCase) aryloxyphenoxyalkanecarboxylic acids
  • HPPD Hydroxyphenylpyruvate dioxygenase
  • ADSL Adenylosuccinate lyase
  • Adenylosuccinate synthase inhibitors of adenylosuccinate synthesis Affected structure / expressed protein characteristic of the plant / tolerance to
  • EPSPS Synthase
  • Protoporphyrinogen oxidase diphenyl ether, cyclic imides, phenylpyrazoles,
  • Cytochrome P450 e.g. P450 SUL Xenobiotics and herbicides such as
  • Antifungal polypeptide AIyAFP Plant pathogens e.g. Septoria and Fusarium
  • Glucose oxidase plant pathogens e.g. Fusarium, Septoria
  • Serine / threonine kinases Plant pathogens, e.g. Fusarium, Septoria and other diseases
  • Double stranded ribonuclease viruses such as BYDV and MSMV
  • Envelope proteins viruses such as BYDV and MSMV
  • Aminopeptidase inhibitors e.g. Leucine Lepidoptera, Coleoptera, Diptera,
  • Lectin Lepidoptera, Coleoptera, Diptera Affected structure / expressed protein characteristic of the plant / tolerance to
  • Protease inhibitors e.g. Cystatin, Patatin, Lepidoptera, Coleoptera, Diptera,
  • Nematodes e.g. Ostrinia nubilalis
  • Heliothis zea armyworms e.g. Spodoptera frugiperda, corn rootworm, Sesamia sp.,
  • Acetolactate synthase (ALS) sulfonylurea compounds imidazolinones
  • Acetyl-CoA-carboxylase (ACCase) aryloxyphenoxyalkanecarboxylic acids
  • HPPD Hydroxyphenylpyruvate dioxygenase
  • ADSL Adenylosuccinate lyase
  • EPSPS Synthase
  • Glyphosate oxidoreductase glyphosate or sulfosate Affected structure / expressed protein characteristic of the plant / tolerance to
  • Protoporphyrinogen oxidase diphenyl ether, cyclic imides,
  • Phenopylate oxadiazoles, etc.
  • Cytochrome P450 e.g. P450 SUL Xenobiotics and herbicides such as
  • Antifungal polypeptide AIyAFP Plant pathogens e.g. Septoria and Fusarium
  • Glucose oxidase plant pathogens e.g. Fusarium, Septoria
  • Serine / threonine kinases Plant pathogens, e.g. Fusarium, Septoria and other diseases
  • Double stranded ribonuclease viruses such as BYDV and MSMV
  • Envelope proteins viruses such as BYDV and MSMV
  • Aminopeptidase inhibitors e.g. Leucine Lepidoptera, Coleoptera, Diptera,
  • Protease inhibitors e.g. Cystatin, Patatin, Lepidoptera, Coleoptera, Diptera,
  • HMG-CoA reductase Lepidoptera, Coleoptera, Diptera Affected structure / expressed protein characteristic of the plant / tolerance to
  • Acetyl-CoA-carboxylase (ACCase) aryloxyphenoxyalkanecarboxylic acids
  • HPPD Hydroxyphenylpyruvate dioxygenase
  • ADSL Adenylosuccinate lyase
  • EPSPS Synthase
  • Protoporphyrinogen oxidase diphenyl ether, cyclic imides,
  • Phenopylate oxadiazoles, etc.
  • Cytochrome P450 e.g. P450 SUL Xenobiotics and herbicides such as
  • Phenylalanine ammonia lyase Plant pathogens, eg bacterial Affected structure / expressed principle Characteristic of the plant / tolerance to
  • Phytoalexins plant pathogens, e.g. bacterial
  • B-1,3-glucanase (antisense) plant pathogens e.g. bacterial
  • Receptor kinase Plant pathogens e.g. bacterial
  • Double stranded ribonuclease viruses such as BYDV and MSMV
  • Envelope proteins viruses such as BYDV and MSMV
  • 3-hydroxysteroid oxidase Lepidoptera e.g. Stem drills, Coleoptera, e.g. Weevils like Lissorhoptrus oryzophilus,
  • Peroxidase Lepidoptera e.g. Stem drills, Coleoptera, e.g. Weevils like Lissorhoptrus oryzophilus,
  • Aminopeptidase inhibitors e.g. Leucine Lepidoptera, e.g. Stem Drill, Coleoptera,
  • Aminopeptidase inhibitor e.g. Weevils such as Lissorhoptrus oryzophilus, Diptera, rice cicadas, e.g. brown-backed rice leafhopper
  • Lectin Lepidoptera eg, borer, Coleoptera
  • Affected structure / expressed principle Characteristic of the plant / tolerance to eg weevils such as Lissorhoptrus oryzophilus, Diptera, rice cicadas, eg brown-backed rice leafhopper
  • Protease inhibitors Lepidoptera, e.g. Stem drills, Coleoptera, e.g. Weevils like Lissorhoptrus oryzophilus,
  • Ribosome inactivating protein Lepidoptera e.g. Stem drills, Coleoptera, e.g. Weevils such as Lissorhoptrus oryzophilus, Diptera, rice cicadas, e.g. brown-backed rice leafhopper
  • HMG-CoA reductase Lepidoptera e.g. Stem drills, Coleoptera, e.g. Weevils such as Lissorhoptrus oryzophilus, Diptera, rice cicadas e.g. brown-backed rice leafhopper
  • Acetolactate synthase (ALS) sulfonylurea compounds imidazolinones
  • Acetyl-CoA-carboxylase (ACCase) aryloxyphenoxyalkanecarboxylic acids
  • HPPD Hydroxyphenylpyruvate dioxygenase
  • ADSL Adenylosuccinate lyase
  • EPSPS Synthase
  • Protopo ⁇ hyrinogen oxidase (PROTOX) diphenyl ether, cyclic imides,
  • Phenopylate oxadiazoles, etc.
  • Cytochrome P450 e.g. P450 SUL or selection of xenobiotics and herbicides such as
  • Antifungal polypeptide AIyAFP Bacterial and fungal pathogens such as
  • Oxalate Oxidase Bacterial and fungal pathogens such as
  • Serine / Threonine Kinases Bacterial and fungal pathogens such as
  • Phenylalanine Ammonia Lyase Bacterial and fungal pathogens such as
  • Phytoalexins plant pathogens e.g. bacterial leaf mildew and rice tan
  • B-1 3-glucanase (antisense) plant pathogens, e.g. bacterial leaf mildew and rice tan
  • Receptor kinase Bacterial and fungal pathogens such as
  • Glucanases Bacterial and fungal pathogens such as Affected structure / expressed principle Characteristic of the plant / tolerance to
  • Double stranded ribonuclease viruses such as BPMV and SbMV
  • Envelope proteins viruses such as BYDV and MSMV
  • Aminopeptidase inhibitors e.g. Leucine lepidoptera, coleoptera, aphids
  • Protease inhibitors e.g. Virgif erin Lepidoptera, Coleoptera, aphids
  • Acetolactate synthase (ALS) sulfonylurea compounds imidazolinones
  • Acetyl-CoA-carboxylase (ACCase) aryloxyphenoxyalkanecarboxylic acids
  • HPPD Hydroxyphenylpyruvate dioxygenase
  • Glutamine synthetase glufosinate, bialaphos Affected structure / expressed protein characteristic of the plant / tolerance to
  • ADSL Adenylosuccinate lyase
  • EPSPS Synthase
  • Protoporphyrinogen oxidase diphenyl ether, cyclic imides,
  • Phenopylate oxadiazoles, etc.
  • Cytochrome P450 e.g. P450 SUL or selection of xenobiotics and herbicides such as
  • Antifungal polypeptide AIyAFP Bacterial and fungal pathogens such as
  • Oxalate Oxidase Bacterial and fungal pathogens such as
  • Serine / Threonine Kinases Bacterial and fungal pathogens such as
  • Cecropin B bacteria such as Corynebacterium sepedonicum, Erwinia carotovora
  • Phenylalanine Ammonia Lyase Bacterial and fungal pathogens such as
  • Receptor kinase Bacterial and fungal pathogens such as
  • Polypeptide with the effect of causing an over bacterial and fungal pathogens such as sensitivity reaction Phytophtora, Verticillium, Rhizoctonia
  • Gene 49 for controlling resistance to bacterial and fungal pathogens such as
  • Glucanases Bacterial and fungal pathogens such as
  • Double-stranded ribonuclease viruses such as PLRV, PVY and TRV
  • Envelope proteins viruses such as PLRV, PVY and TRV
  • 17 kDa or 60 kDa protein viruses such as PLRV, PVY and TRV
  • Inclusion proteins of the nucleus e.g. a or b viruses such as PLRV, PVY and TRV
  • Pseudoubiquitin viruses such as PLRV, PVY and TRV
  • Replicase viruses such as PLRV, PVY and TRV
  • 3-hydroxysteroid oxidase Coleoptera e.g. Colorado beetle, aphids
  • Peroxidase Coleoptera e.g. Colorado beetle, aphids
  • Aminopeptidase inhibitors e.g. Leucine Coleoptera, e.g. Colorado beetle, aphids
  • Stilbene synthase Coleoptera e.g. Colorado beetle, aphids
  • Lectins Coleoptera e.g. Colorado beetle, aphids
  • Protease inhibitors e.g. Cystatin, Patatin Coleoptera, e.g. Colorado beetle, aphids
  • Ribosome inactivating protein Coleoptera e.g. Colorado beetle, aphids
  • HMG-CoA-Reductase Coleoptera eg Colorado beetle, aphids Affected structure / expressed protein characteristic of the plant / tolerance to
  • Acetolactate synthase (ALS) sulfonylurea compounds imidazolinones
  • Acetyl-CoA-carboxylase (ACCase) aryloxyphenoxyalkanecarboxylic acids
  • HPPD Hydroxyphenylpyruvate dioxygenase
  • ADSL Adenylosuccinate lyase
  • EPSPS Synthase
  • Protoporphyrinogen oxidase diphenyl ether, cyclic imides,
  • Phenopylate oxadiazoles, etc.
  • Cytochrome P450 eg P450 SUL or selection of xenobiotics and herbicides such as Affected structure / expressed principle Characteristic of the plant / tolerance to
  • Antifungal polypeptide AIyAFP Bacterial and fungal pathogens such as bacterial spot disease, Fusarium,
  • Pyrrolnitrin Synthesis Genes Bacterial and fungal pathogens such as bacterial spot disease, Fusarium,
  • Serine / Threonine Kinases Bacterial and fungal pathogens such as bacterial spot disease, Fusarium,
  • Cecropin B Bacterial and fungal pathogens such as bacterial spot disease, Fusarium,
  • Phenylalanine ammonia lyase Bacterial and fungal pathogens such as bacterial spot disease, Fusarium,
  • Bacterial and fungal pathogens such as bacterial spot disease, Fusarium,
  • Polygalacturonase inhibitors Bacterial and fungal pathogens such as bacterial spot disease, Fusarium,
  • Receptor kinase Bacterial and fungal pathogens such as bacterial spot disease, Fusarium,
  • Chitinases Bacterial and fungal pathogens such as bacterial spot disease, Fusarium, Affected structure / expressed principle Characteristic of the plant / tolerance to
  • Bamase Bacterial and fungal pathogens such as the bacterial spot disease, Fusarium,
  • Glucanases Bacterial and fungal pathogens such as bacterial spot disease, Fusarium,
  • Double stranded ribonuclease viruses such as PLRV, PVY and ToMoV
  • Envelope proteins viruses such as PLRV, PVY and ToMoV
  • 17 kDa or 60 kDa protein viruses such as PLRV, PVY and ToMoV
  • Inclusion proteins of the nucleus e.g. a or b viruses such as PLRV, PVY and ToMoV or
  • Pseudoubiquitin viruses such as PLRV, PVY and ToMoV
  • Replicase viruses such as PLRV, PVY and ToMoV
  • 3-hydroxysteroid oxidase Lepidoptera e.g. Heliothis, white fly
  • Peroxidase Lepidoptera e.g. Heliothis, white fly
  • Aminopeptidase inhibitors e.g. Leucine Lepidoptera e.g. Heliothis, white fly,
  • Lectin Lepidoptera e.g. Heliothis, white fly
  • Protease inhibitors e.g. Cystatin, patatin Lepidoptera e.g. Heliothis, white fly,
  • Ribosome inactivating protein Lepidoptera e.g. Heliothis, white fly
  • Stilbene synthase Lepidoptera eg Heliothis, white fly, Affected structure / expressed principle Characteristic of the plant / tolerance to
  • HMG-CoA reductase Lepidoptera e.g. Heliothis, white fly
  • Acetolactate synthase (ALS) sulfonylurea compounds imidazolinones
  • Acetyl-CoA-carboxylase (ACCase) aryloxyphenoxyalkanecarboxylic acids
  • HPPD Hydroxyphenylpyruvate dioxygenase
  • ADSL Adenylosuccinate lyase
  • EPSPS Synthase
  • Protoporphyrinogen oxidase (PROTOX) diphenyl ether, cyclic imides, Affected structure / expressed protein characteristic of the plant / tolerance to
  • Phenopylate oxadiazoles, etc.
  • Cytochrome P450 e.g. P450 SUL or selection of xenobiotics and herbicides such as
  • Cecropin B Bacterial and fungal pathogens, rot,
  • Phenylalanine Ammonia Lyase Bacterial and fungal pathogens
  • Cf genes e.g. Cf 9 Ct5 Cf4 Cf2 Bacterial and fungal pathogens
  • Double-stranded ribonuclease viruses such as CMV, TEV
  • Envelope proteins viruses such as CMV, TEV Affected structure / expressed protein characteristic of the plant / tolerance to
  • Inclusion proteins of the nucleus e.g. a or b viruses such as CMV, TEV or
  • Pseudoubiquitin viruses such as CMV, TEV
  • Replicase viruses such as CMV, TEV
  • Aminopeptidase inhibitors e.g. Leucin lepidoptera, white fly, aphids
  • Lectin Lepidoptera white fly, aphids
  • Protease inhibitors e.g. Cystatin, Patatin Lepidoptera, white fly, aphids
  • Stilbene synthase Lepidoptera white fly, aphids
  • HMG-CoA reductase Lepidoptera white fly, aphids
  • Acetolactate synthase (ALS) sulfonylurea compounds imidazolinones
  • Acetyl-CoA-carboxylase (ACCase) aryloxyphenoxyalkanecarboxylic acids
  • HPPD Hydroxyphenylpyruvate dioxygenase
  • ADSL Adenylosuccinate lyase
  • EPSPS Synthase
  • Protoporphyrinogen oxidase diphenyl ether, cyclic imides,
  • Phenopylate oxadiazoles, etc.
  • Cytochrome P450 e.g. P450 SUL or selection of xenobiotics and herbicides such as
  • Oxalate Oxidase Bacterial and fungal pathogens such as
  • Serine / Threonine Kinases Bacterial and fungal pathogens such as
  • Cecropin B Bacterial and fungal pathogens like Affected structure / expressed principle Characteristic of the plant / tolerance to
  • Phenylalanine Ammonia Lyase Bacterial and fungal pathogens such as
  • Cf genes e.g. Cf 9 Cf5 Cf4 Cf2 Bacterial and fungal pathogens such as
  • Receptor kinase Bacterial and fungal pathogens such as
  • Glucanases Bacterial and fungal pathogens such as botrytis and powdery mildew
  • 17kDa or 60 kDa protein viruses Affected structure / expressed principle Characteristic of the plant / tolerance to
  • Inclusion proteins of the nucleus e.g. a or b viruses or
  • Aminopeptidase inhibitors e.g. Leucine lepidoptera, aphids
  • Protease inhibitors e.g. Cystatin, Patatin Lepidoptera, aphids
  • Acetolactate synthase (ALS) sulfonylurea compounds imidazolinones
  • Acetyl-CoA-carboxylase (ACCase) aryloxyphenoxyalkanecarboxylic acids
  • HPPD Hydroxyphenylpyruvate dioxygenase
  • ADSL Adenylosuccinate lyase
  • EPSPS Synthase
  • Protoporphyrinogen oxidase diphenyl ether, cyclic imides,
  • Phenopylate oxadiazoles, etc.
  • Cytochrome P450 e.g. P450 SUL or selection of xenobiotics and herbicides such as
  • Oxidase (antisense) Cylindrosporium, Phoma, Sclerotinia

Landscapes

  • Life Sciences & Earth Sciences (AREA)
  • Agronomy & Crop Science (AREA)
  • Pest Control & Pesticides (AREA)
  • Plant Pathology (AREA)
  • Health & Medical Sciences (AREA)
  • Engineering & Computer Science (AREA)
  • Dentistry (AREA)
  • General Health & Medical Sciences (AREA)
  • Wood Science & Technology (AREA)
  • Zoology (AREA)
  • Environmental Sciences (AREA)
  • Agricultural Chemicals And Associated Chemicals (AREA)
  • Breeding Of Plants And Reproduction By Means Of Culturing (AREA)
EP07856774A 2006-12-27 2007-12-15 Verfahren zur verbesserten nutzung des produktionspotentials transgener pflanzen Withdrawn EP2104423A1 (de)

Applications Claiming Priority (3)

Application Number Priority Date Filing Date Title
DE102006061621 2006-12-27
DE102007009957A DE102007009957A1 (de) 2006-12-27 2007-03-01 Verfahren zur verbesserten Nutzung des Produktionsptentials transgener Pflanzen
PCT/EP2007/011037 WO2008080545A1 (de) 2006-12-27 2007-12-15 Verfahren zur verbesserten nutzung des produktionspotentials transgener pflanzen

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EP2104423A1 true EP2104423A1 (de) 2009-09-30

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US (1) US20100313310A1 (pt)
EP (1) EP2104423A1 (pt)
JP (1) JP2010514714A (pt)
KR (1) KR20090105949A (pt)
AR (1) AR064560A1 (pt)
AU (1) AU2007341651B2 (pt)
BR (1) BRPI0720712A2 (pt)
CL (1) CL2007003844A1 (pt)
DE (1) DE102007009957A1 (pt)
MX (1) MX2009006943A (pt)
TW (2) TWI549609B (pt)
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AU2007341651A1 (en) 2008-07-10
MX2009006943A (es) 2009-07-10
US20100313310A1 (en) 2010-12-09
AU2007341651B2 (en) 2014-03-20
JP2010514714A (ja) 2010-05-06
TWI549609B (zh) 2016-09-21
TW200843638A (en) 2008-11-16
TW201417709A (zh) 2014-05-16
KR20090105949A (ko) 2009-10-07
CL2007003844A1 (es) 2008-05-16
DE102007009957A1 (de) 2008-07-03
AR064560A1 (es) 2009-04-08
BRPI0720712A2 (pt) 2014-02-18

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